The Tissue System
Based on structure and location, tissue systems are of three types:
- Epidermal tissue system
- Ground (fundamental) tissue system
- Vascular (conducting) tissue system
1. Epidermal Tissue System
- Forms the outermost covering of the whole plant body.
- Comprises epidermal cells, stomata, and epidermal appendages (trichomes & hairs).
Epidermis
- The outermost layer of the primary plant body.
- Usually single-layered.
- Made up of elongated, compactly arranged parenchymatous cells with a small amount of cytoplasm lining the cell wall and a large vacuole.
- The outside of the epidermis is often covered with a waxy thick layer (cuticle) to prevent water loss. The cuticle is absent in roots.
Stomata
- Structures present in the epidermis of leaves.
- Regulate transpiration and gaseous exchange.
- A stoma is made of two bean-shaped cells (guard cells).
- In grasses, guard cells are dumbbell-shaped.
- The outer walls of guard cells (away from the stomatal pore) are thin, and the inner walls (towards the stomatal pore) are highly thickened.
- Guard cells possess chloroplasts and regulate the opening and closing of stomata.
- Some epidermal cells near the guard cells may become specialized in shape and size, called subsidiary cells.
- The stomatal aperture, guard cells, and surrounding subsidiary cells together form the stomatal apparatus.
Epidermal Appendages
- Root hairs: Unicellular elongations of epidermal cells that absorb water and minerals from the soil.
- Trichomes: Epidermal hairs on the stem, usually multicellular, branched or unbranched, soft or stiff, and sometimes secretory. They help prevent water loss due to transpiration.
2. The Ground Tissue System
- All tissues except epidermis and vascular bundles constitute the ground tissue.
- Consists of simple tissues (parenchyma, collenchyma, and sclerenchyma).
- Parenchymatous cells are present in cortex, pericycle, pith, and medullary rays in primary stems and roots.
- In leaves, the ground tissue consists of thin-walled chloroplast-containing cells, called mesophyll.
3. The Vascular Tissue System
- Consists of complex tissues (xylem and phloem).
- Xylem and phloem together constitute vascular bundles.
Based on the presence or absence of cambium, vascular bundles are of two types:
- Open type: Cambium is present between phloem and xylem, allowing the formation of secondary xylem and phloem tissues. Example: dicotyledonous stems.
- Closed type: Cambium is absent, so secondary tissues are not formed. Example: monocotyledons.
Based on the arrangement of xylem and phloem, vascular bundles are of two types:
- Radial type: Xylem and phloem are arranged alternately on different radii. Seen in roots.
- Conjoint type: Xylem and phloem are situated at the same radius. Seen in stems and leaves. Conjoint vascular bundles usually have phloem located only on the outer side of xylem.

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